Zanthoxylum hawaiiense

Hillebr. (1888)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Zanthoxylum

aʻe [ae], hea‘e, mānele, zanthoxylum

Description

Key Characters:

Growth Form: Small to medium-sized trees (1–)3–8 m tall.

Stems: Trunk up to 25 cm in diameter, bark pale to dark gray, foliage usually lemon–scented, new growth glabrous or finely puberulent or tomentose, young branches subtended by a pair of bud scales.

Roots:

Leaves: Leaves compound. Alternate. Blades broadly deltate–ovate or lanceolate, when deltate–ovate, the lateral ones asymmetrical; leaflets 3, thin to sometimes thick, coriaceous, 3.4–10(–12) cm long, 1.5–5(–7.5) cm wide, lateral petiolules articulated only at base, ± thickened near leaflet but not actually articulated, (3–)10–40 mm long, terminal petiolule twice articulated, at base and near leaflet, (12–)15–52 mm long. Apex acute to acuminate, sometimes slightly falcate. Base ± asymmetrically truncate or cuneate. Surfaces glabrous or lower surfaces finely puberulent, glandular punctate, the glands scattered and prominent. Margins minutely crenulate. Lateral veins 5–9 pairs, strongly ascending, higher order venation forming a reticulate pattern. Rachis terete (cylindrical) or winged. Stipules absent.

Flowers: Flowers usually 15–20 in axillary, open, cymose inflorescences usually 4–8 cm long, peduncles 20–50 mm long, pedicels 2–4 mm long; bracteoles ca. 1 mm long, puberulent. Flowers unisexual, actinomorphic. Calyx of 4 sepals, alternate with the petals, narrowly deltate, ca. 1 mm long, sparsely puberulent and ciliolate. Corolla of 4 petals, color unknown, oblong–lanceolate to narrowly ovate, 3–6 mm long. Stamens 4, glabrous, absent in pistillate flowers(?); anthers medifixed, dithecal. Ovary superior, 1-carpellate; ovule placentation axile, rudimentary in staminate flowers; style oblique; stigma capitate.

Fruit: Follicles 1; falcate; obovoid; 8–10 mm long; apex rounded; exocarp rugulose and pitted. Seeds 1 per fruit; on a slender funicle; black; subglobose; compressed; 7–8 mm in diameter.

Ploidy:

Habitat: Dry forest on lava flows; sometimes in mesic forest.

Elevation Range: 550–1,740 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Fl. Hawaiian Isl.: 76 (1888)

Other References

Wagner et al. 1990:1214 (Mo, L, M, H); Lorence et al. 1995:54 (K); Wood 2007:16 (WM); Hank Oppenheimer pers. obs./Forbes 2.L BISH (L†, 1913)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Vegetation: Diospyros, Nestegis, Metrosideros, Pleomele, Pittosporum, Dodonaea, Claoxylon, Antidesma, Psychotria, Sophora, Psydrax, Grevillea, Psidium, Osteomeles, Microlepia, Alyxia, Doodia, Dryopteris, Achyranthes, Dubantia. Preserved_Specimen Oppenheimer, H.L. 91306 Maui BISH 2013-09-11
2 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details On dry a`a lava in open, native forest. Preserved_Specimen Herbst, D.R. 730 Maui BISH 1967-11-12
3 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Not common on Aa flows dominated by Metrosideros and Myrsine lanaiense. Preserved_Specimen Marr, K.L. 93 Hawaii BISH 1988-07-31
4 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Not common on Aa flows dominated by Metrosideros and Myrsine lanaiense. Preserved_Specimen Marr, K.L. 35 Hawaii BISH 1988-06-02
5 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Dry ohia forest with mixed native trees and a native shrub-exotic grass-shrub u/s. Full sun; in small depression. Rare; growing in sm depression on pahoehoe. Preserved_Specimen Jacobi, J.D. 1009 Hawaii BISH 1976-01-01
6 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Lava flows. Preserved_Specimen Rock, J.F.C. Maui BISH 1954-02-01
7 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Moderately dry sunny canyon. Preserved_Specimen Degener, O. 8604 Molokai BISH 1928-05-21
8 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details In Dodonaea-Styphelia lowland mesic shrubland with Vaccinium, Neraudia, Lipochaeta, Bidens, Myoporum, Wikstroemia, Metrosideros, Pleomele, Pittosporum, Chamaesyce, Lysimachia, Viola, Pipturus, Urera, Lantana, Melinis, Schiedea. Preserved_Specimen Perlman, S. 6678 Molokai BISH 1987-10-23
9 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Metrosideros-Acacia montane mixed mesic forest, summer dry Diospyros sandwicensis, Pleomele aurea, Myrsine lanaiensis, Planchonella sandwicensis, Cryptocarya mannii, Myoporum sandwicensis, Nestegis sandwicensis, Psydrax odorata, Zanthoxylum dipetalum, Pteraly Preserved_Specimen Wood, K.R. 15417 Kauai BISH 2013-02-17
10 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Not common on Aa flows dominated by Metrosideros and Myrsine lanaiense. Preserved_Specimen Marr, K.L. 88 Hawaii BISH 1988-07-31
11 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Growing on steep, rocky SE side of valley. Associated vegetation: Metrosideros, Nestegis, Diospyros, Pouteria, Remya, Zanthoxylum, Neraudia, Myrsine, Melicope, Cocculus, Sadleria, Pleomele, Dubautia, Coprosma, Alyxia, Charpentiera, Peperomia, Doodia. Preserved_Specimen Oppenheimer, H.L. 60625 Maui BISH 2006-06-21
12 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details In open south facing slope in grazed area; on a`a substrate, covered with kikuyu grass. Vegetation: Degraded Dry/Mesic Forest remnant, with Nestegis, Streblus, Ochrosia, Pleomele, Myrsine, Dodonaea, Santalum, Chamaesyce, Diospyros, Osteomeles, Cocculus, Alyxia, Asplenium, Lepisorus, Peperomia, Plectranthus, Alectryon, Cyperus, Bocconia, Lantana, Schinus, Cinnamomum, Pennisetum, Bidens, Passiflora, Neonotonia, Senecio, Melinis, Gomphrena Preserved_Specimen Oppenheimer, H.L. 90905 Maui BISH 2009-09-18
13 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details On lava flows. Preserved_Specimen Rock, J.F.C. 25719 Hawaii BISH 1957-03-14
14 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Ravine, smothered with Lantana with isolated trees poking out. Assoc. trees: Melia, Metrosideros, Dodonaea, Alectryon, Antidesma, Pisonia & Charpentiera. Mesic-forested area. Preserved_Specimen Lau, J. 2204 Kauai BISH 1986-03-30
15 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Not common on Aa flows dominated by Metrosideros and Myrsine lanaiense. Preserved_Specimen Marr, K.L. 89 Hawaii BISH 1988-07-31
16 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details mixed mesic shrubland, grassland, steep to vertical rocky slopes and ridges; Leptecophylla tameiameiae, Dodonaea viscosa, Dianella sandwicensis, Morelotia gahniiformis, Dicranopteris linearis, Bidens micrantha, Neraudia melastomifolia, Melicope hawaiiense, Zanthoxylum hawaiiensis Preserved_Specimen Wood, K.R. 11926 Maui BISH 2006-06-20
17 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details small drainage, mixed mesic forest; Metrosideros polymorpha, Cheirodendron trigynum, Dodonaea viscosa, Nestegis sandwicensis, Hedyotis acuminata, Cyrtandra grayi, Melicope hawaiiense Preserved_Specimen Wood, K.R. 11940 Maui BISH 2006-06-21
18 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Diverse mesic forest; w/ Bobea timonioides, Melicope haupuensis, M. knudsenii, Alectryon, Peucedanum, Pteralyxia. North aspect. Preserved_Specimen Wood, K.R. 2127 Kauai BISH 1993-11-12
19 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Montane mixed dry to mesic forest, just below Metrosideros-Acacia forest with Diospyros sandwicensis, Chrysodracon cf. stenophylla (=aurea), Myrsine lanaiensis, Planchonella sandwicensis, Cryptocarya mannii, Myoporum sandwicensis, Nestegis sandwicensis, Psydrax odorata, Zant Preserved_Specimen Wood, K.R. 16472 Kauai BISH 2015-06-04
20 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Wet woods. Preserved_Specimen St.John, H. 23462 Molokai BISH 1948-12-22
21 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Old `a`a lava. Preserved_Specimen MacDaniels, L.H. 311 Hawaii BISH 1926-12-01
22 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Vegetation: Open Metrosideros forest. Growing on aa lava, either by its self of growing in with Metrosideros trees. Preserved_Specimen Warshauer, F.R. 815 Hawaii BISH 1977-01-11
23 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Beside cliff, with Charpentiera Preserved_Specimen Wood, K.R. 289 Kauai BISH 1990-03-31
24 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Not common on Aa flows dominated by Metrosideros and Myrsine lanaiense. Preserved_Specimen Marr, K.L. 94 Hawaii BISH 1988-07-31
25 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Mesic ohia forest with other native trees and native shrub-exotic grass and shrub u/s. Preserved_Specimen Warshauer, F.R. 3021 Molokai BISH 1980-08-07
26 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details On steep SE side of valley. Associated vegetation: Metrosideros, Dodonaea, Myrsine, Coprosma, Wikstroemia, Vaccinium, Sophora, Melicope, Remya, Pteridium, Microlepia, Dubautia, Leptecophylla, Lysimachia, Pteris, Pteridium, Dryopteris. Threats: Rockslides, Buddleia asiatica, Ageratina adenophora. Preserved_Specimen Oppenheimer, H.L. 60620 Maui BISH 2006-06-20
27 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Dry `ohi`a forest with mixed native trees and a native shrub and mixed grass understory. Preserved_Specimen Warshauer, F.R. 1877 Hawaii BISH 1978-06-25
28 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details One tree seen, in sparse scrub forest on rough lava. Preserved_Specimen Fosberg, F.R. 60612 Hawaii BISH 1980-09-10
29 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Diospyros sandwicensis-Metrosideros mesic forest. With Pleomele auwahiensis, Myrisine lanaiensis, Streblus, Nothocestrum latifolium, Pittosporum argentifolium, Neraudia sericea, Zanthoxylum hawaiiensis, Ochrosia compta, Schinus terebinthifolius, Melinus Preserved_Specimen Perlman, S. 22295 Molokai BISH 2009-12-14
30 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details In Dodonaea-Styphelia lowland mesic shrubland with Vaccinium, Neraudia, Lipochaeta, Bidens, Myoporum, Wikstroemia, Metrosideros, Pleomele, Pittosporum, Chamaesyce, Lysimachia, Viola, Pipturus, Urera, Lantana, Melinis, Schiedea. Preserved_Specimen Perlman, S. 6678 Molokai BISH 1987-10-23
31 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details `A`a lava. Preserved_Specimen Rock, J.F.C. Hawaii BISH 1912-03-01
32 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Two trees observed at this location. Vegetation: Myrsine, Coprosma, Urera, Pouteria, Wikstroemia, Dodonaea, Cyrtandra, Metrosideros, Alyxia, Remya, Peperomia, Asplenium, Cyrtomium. Preserved_Specimen Oppenheimer, H.L. 80636 Maui BISH 2006-08-23
33 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1881 Maui BISH 1920-03-07
34 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1926 Maui BISH 1920-03-11
35 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1962 Maui BISH 1920-03-14
36 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 8658 Maui BISH 1910-11-01
37 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 10205 Hawaii BISH 1912-03-01
38 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 124 Molokai BISH 1912-07-01
39 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 8657 Maui BISH 1910-11-01
40 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 2068 Maui BISH 1920-03-23
41 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1926 Maui BISH 1920-03-11
42 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1881 Maui BISH 1920-03-07
43 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Faurie, A. 250 Molokai BISH 1910-06-01
44 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1956 Maui BISH 1920-03-17
45 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 10204 Hawaii BISH 1912-03-01
46 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 8076 Lanai BISH 1910-07-24
47 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 124 Molokai BISH 1912-07-01
48 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 10204 Hawaii BISH 1912-03-01
49 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1952 Maui BISH 1920-03-13
50 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 124 Molokai BISH 1912-07-01
51 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Degener, O. 26868 Maui BISH 1960-01-17
52 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 27007 Maui BISH 1953-02-01
53 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 124 Molokai BISH 1912-07-01
54 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 240 Molokai BISH 1912-07-01
55 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1956 Maui BISH 1920-03-17
56 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 1881 Maui BISH 1920-03-07
57 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Degener, O. 26868 Maui BISH 1960-01-17
58 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 124 Molokai BISH 1912-07-01
59 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Pratt, T.K. 87 Molokai BISH 1972-01-13
60 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. Maui BISH 1920-03-01
61 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Wilder, G.P. Maui BISH 1913-01-01
62 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Wilder, G.P. Maui BISH 1913-01-01
63 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. Maui BISH 1920-03-01
64 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 5207 BISH 1909-09-18
65 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Greenwell, A.B.H. 19803 Hawaii BISH 1949-08-21
66 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 124 Molokai BISH 1912-07-01
67 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Forbes, C.N. 2 Lanai BISH 1913-06-01
68 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 8373 Hawaii BISH 1910-06-01
69 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 5207 Kauai BISH 1909-09-18
70 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Rock, J.F.C. 10204 Hawaii BISH 1912-03-01
71 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hillebrand, W.B. Hawaii BISH
72 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hillebrand, W.B. Lanai BISH
73 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hillebrand, W.B. Hawaii BISH
74 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hillebrand, W.B. Lanai BISH
75 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Hillebrand, W.B. Hawaii BISH
76 Zanthoxylum hawaiiense Locality redacted. Contact Bishop Museum Botany Department for details Metrosideros-Diospyros forest. With Nestegis sandwicensis, Sophora chrysophylla, Pyschotria kaduana, Lipochaeta connata, Remya mauiensis, Pleomele, Dodonaea viscosa, Pisonia Preserved_Specimen Perlman, S. 22831 Maui BISH 2012-04-17